Kappes PJ, KM Dugger, A Lescroël, DG Ainley, G Ballard, KJ Barton, PO’B Lyver, PR Wilson. 2021. Age-related reproductive performance of the Adélie Penguin, a long-lived seabird exhibiting similar outcomes regardless of individual life-history strategy. Journal of Animal Ecology 2021;90:931-942, https://doi.org/10.1111/1365-2656.13422.

Anthony, CR, CA Hagen, KM Dugger, and RD Elmore. 2021. Greater Sage-Grouse nest bowls buffer microclimate in a post-megafire landscape although effects on nest survival are marginal. Ornithological Applications 123:1-13. https://doi.org/10.1093/ornithapp/duaa068.

AB Franklin, KM Dugger, DB Lesmeister, RJ Davis, JD Wiens, GC White, JD Nichols, JE Hines, CB Yackulic, CJ Schwarz, SH Ackers, LS Andrews, LL Bailey, R Bown, J Burgher, KP Burnham, PC Carlson, T Chestnut, MM Conner, KE Dilione, ED Forsman, EM Glenn, SA Gremel, KA Hamm, DR Herter, JM Higley, RB Horn, JM Jenkins, WL Kendall, DW Lamphear, C McCafferty, TL McDonald, JA Reid, JT Rockweit, DC Simon, SG Sovern, JK Swingle, and H Wise. 2021. Range-wide declines of Northern Spotted Owls populations in the Pacific Northwest: a meta-analysis. Biological Conservation Volume 259, DOI: 10.1016/j.biocon.2021.109168.

Morandini, V, KM Dugger, A Lescroël, A Schmidt and G Ballard. 2021. Maintenance of nest quality in Adélie penguins: an additional benefit to life in the center. Polar Biology 44:1553-11562, DOI://doi.org/10.1007/s00300-021-02894-5.

Wiens, JD, KM Dugger, M Higley, DB Lesmeister, AB Franklin, KA Hamm, GC While, KE Dilione, DC Simon, RR Bown, PC Carlson, CB Yackulic, JD Nichols, JE Hines, RJ Davis, DW Lamphear, C McCafferty, TL McDonald, and SG Sovern. 2021. Invader removal triggers competitive release in a threatened avian predator. Proceedings of the National Academy of Science 118 (31): e2102859118. https://doi.org/10.1073/pnas.2102859118.

Jenkins JMA, DB Lesmeister, ED Forsman, KM Dugger, SH Ackers, LS Andrews, SA Gremel, B Hollen, CE McCafferty, MS Pruett, JA Reid, SG Sovern, and JD Wiens. 2021. Conspecific and congeneric interactions shape increasing rates of breeding dispersal of northern spotted owls. Ecological Applications e02398, http://doi.org/10.1002/eap.2398.

Schmidt, AE, G Ballard, A Lescroël, KM Dugger, D Jongsomjit, ML Elrod, and DG Ainley. 2021. The influence of subcolony-scale nesting habitat on reproductive success of Adélie penguins. Scientific Reports 11:15380, https://doi.org/10.1038/s41598-021-94861-7.


Marks-Fife, CA, ED Forsman, and KM Dugger. 2020. Age structure of red tree voles in spotted owl pellets estimated with regression models of molar tooth development. Northwest Science 93:193-208, https://doi.org/10.3955/046.093.0304.

Anthony, CR, CA Hagen, KM Dugger, and RD Elmore. 2020. The effects of fire on the thermal environment of sagebrush communities. Journal of Thermal Biology 89:102488. https://doi.org/10.1016/j.jtherbio.2019.102488.

Lescroël, A, P O’B Lyver, D Jongsomjit, S Veloz, KM Dugger, P Kappes, BJ Karl, AL Whitehead, R Pech, TL Cole, G Ballard. 2020. Individual quality and sex are the main drivers of inter-individual differences in the foraging behavior of breeding Adélie penguins. Marine Ecology Progress Series 636: 189–205, DOI: /10.3354/meps13208.

Duchac, LS, DB Lesmeister, KM Dugger, ZJ Ruff, and RJ Davis. 2020. Passive acoustic monitoring effectively detects Northern Spotted Owls and Barred Owls over a range of forest conditions. The Condor 122: 1–22, DOI: 10.1093/condor/duaa017.

Morandini, V, KM Dugger, DG Ainley, and M Ferrer. 2020. Rockhopper Penguin-Imperial Cormorant mixed colonies in the Falkland Islands: a stroke of luck for late breeders. Eosphere 11(11):e03272. doi: 10.1002/ecs2.3272.


Foster, LJ, KM Dugger, CA Hagen, and DA Budeau. 2019. Acute Effects of Wildfire on Greater Sage-Grouse Vital Rates. Journal of Wildlife Management 83:121–134; 2019; DOI: 10.1002/jwmg.21573.

Sovern, SG, DB Lesmeister, KM Dugger, MS Pruett, RJ Davis, and JM Jenkins. 2019. Activity Center Selection by Northern Spotted Owls. Journal of Wildlife Management 83:714-727; https://doi.org/10.1002/jwmg.21632.

Lescroël, A, G Ballard, M Massaro, KM Dugger, S Jennings, A Pollard, E Porzig, A Schmidt, A Varsani, D Grémillet, and D Ainley. 2019. Evidence of age-related improvement in the foraging efficiency of Adélie penguins. Scientific Reports 9:3375, DOI:/10.1038/s41598-019-39814-x.

Yackulic, CB, LL Bailey, KM Dugger, RJ Davis, AB Franklin, ED Forsman, SH Ackers, LS Andrews, LV Diller, SA Gremel, KA, Hamm, DR Herter, JM Higley, RB Horn, C McCafferty, JA Reid, JT Rockweit, and SG Sovern. 2019. The past and future roles of competition and habitat in the range-wide occupancy dynamics of northern spotted owls: Ecological Applications, 29(3) e01861, doi.org/10.1002/eap.1861.

Morandini, V, A Lescröel, D Jongsomjit, S Winquist, A Schmidt, G Ballard, P Kappes, and KM Dugger. 2019. Prevalence of three-chick nests in Adelie penguin Pygoscelis Adeliae at Cape Crozier, Ross Island. Marine Ornithology 47:77-80.

Wiens, JD, KM Dugger, DB Lesmeister, KE Dilione, and DC Simon. 2019. Effects of Barred Owl (Strix varia) removal on population demography of Northern Spotted Owls in Washington and Oregon, 2015-18. U.S. Geological Survey Open File Report 2019-1074, 17pp. hppts://doi.org./10.3133/ofr20191074.

Mangan, AO, T Chestnut, JC Vogeler, IK Breckheimer, WM King, KE Bagnall, and KM Dugger. 2019. The effects of habitat, weather and Barred Owls (Strix varia) on breeding and occupancy dynamics of Northern Spotted Owls (S. occidentalis caurina). Condor 121:1-20, DOI: 10.1093/condor/duz031.

Morandini, V, KM Dugger, G Ballard, M Elrod, A Schmidt, V Ruoppolo, A Lescroël, D Jongsomjit, M Massaro, J Pennycook, GL Kooyman, K Schmidlin, S Kraberger, D Ainley, and A Varsani. 2019. Identification of a novel Adélie penguin circovirus at Cape Crozier (Ross Island, Antarctica). Viruses 11, 1088; doi:10.3390/v11121088.

Jenkins, JMA, DB. Lesmeister, ED Forsman, KM Dugger, SH Ackers, LS Andrews, CE McCafferty, MS Pruett, JA Reid, SG Sovern, RB Horn, SA Gremel, JD Wiens, and Z Yang. 2019. Social status, forest disturbance, and barred owls shape long-term trends in breeding dispersal distance of Northern Spotted Owls. The Condor 121:1–17, DOI: 10.1093/condor/duz055.


Gremillet, D., A. Lescroel, G. Ballard, K.M. Dugger, M. Massaro, and D.G. Ainley. 2018. Energetic fitness: confronting field metabolic rates assessed via accelerometry with conventional fitness metrics. Functional Ecology 2018;1–11, DOI: 10.1111/1365-2435.13074

Miller, MP, SM Haig, ED Forsman, RG Anthony, L Diller, KM Dugger, AB Franklin, SA Gremel, DB Lesmeister, M Higley, DR Herter, and SG Sovern. 2018. Variation in inbreeding rates across the range of Northern Spotted Owls (Strix occidentalis caurina)- insights from over 30 years of monitoring data: The Auk: Ornithological Advances, v. 135, no. 4, p. 821-833, https://doi.org/10.1642/AUK-18-1.1.

Ainley, DG, KM Dugger, M La Mesa, G Ballard, KJ Barton, S Jennings, BJ Karl, A Lescroël, PO’B Lyver, A Schmidt, and P Wilson. 2018. Post-fledging survival of Adélie penguins at multiple colonies: Chicks raised on fish do well. Marine Ecology Progress Series 601:239-251, DOI: 10.3354/meps12687.

Schuyler, EM, KM Dugger, and DH Jackson. 2018. Effects of Distribution, Behavior, and Climate on Mule Deer Survival. Journal of Wildlife Management 83:89–99, DOI: 10.1002/jwmg.21558.

Foster, LJ, KM Dugger, CA Hagen and DA Budeau. 2018. Potential Effects of GPS Transmitters on Greater Sage-Grouse Survival in a Post-Fire Landscape. Wildlife Biology 2018: wlb.00479, doi: 10.2981/wlb.00479. [IP-094455, approved 7/5/2018] [C:50, D:0, I:60, W:35].


Van Doorslaer, K, V. Ruoppolo, A. Schmidt, A. Lescroel, D. Jongsomjit, M. Elrod, S. Kraberger, D. Stainton, K.M. Dugger, G. Ballard, D.G. Ainley, and A. Varsani. 2017. Evidence of genome plasticity in non-mammalian papillomaviruses.  Virus Evolution 3(2): vex027. doi: 10.1093/ve/vex027

Youngflesh, C., S. Jenouvrier, Y. Li, R. Ji, D.G., Ainley, G. Ballard, C. Barbraud, K. Delord, K. M. Dugger, L. M. Emmerson, W. R. Frazer, J. T. Hinke, P. O’B. Lyver, S. Omastroni, C. J. Southwell, S. G. Trivelpiece, W. Z. Trivelpiece, and H. J. Lynch.  2017.  Circumpolar analysis of the Adélie Penguin reveals the importance of environmental variability in phenological mismatch. Ecology 98: 940-951.


Krementz, D. G., KL Willard, JM Carroll, and KM Dugger.  2016.  King rail (Rallus elegans) nesting and brood-rearing ecology in a managed wetland in Oklahoma, USA.  Waterbirds 39:241-249.

Wilson, DJ, PO’B Lyver, TC Greene, AL Whitehead, KM Dugger, BJ Karl, JRF Barringer, R McGarry, AM Pollard, and DG Ainley. 2016. South polar skua breeding populations in the Ross Sea assessed from demonstrated relationship with Adélie penguin numbers. Polar Biology, Online first: DOI 10.1007/s00300-016-1980-4.

Wallace, ZP, PL Kennedy, JR Squires, RJ Oakleaf, KM Dugger, and LE Olson. 2016. Re-occupancy of breeding territories by ferruginous hawks in Wyoming: relationships to environmental and anthropogenic factors. PLoS ONE 11(4): 0152977. doi:10.1371/journal.pone.0152977

Jennings, S, A Varsani, KM Dugger, G Ballard, and DG Ainley. 2016. Sex-based differences in Adélie penguin (Pygoscelis adeliae) chick growth rates. PloS ONE 11(3): e0149090. doi:10.1371

Dugger, BD, TT Fox, JM Coluccy, KM Dugger, MJ Petrie, and D Kraege. 2016. Mallards in Eastern Washington: breeding demographics for a species outside its core distribution. Journal of Wildlife Management 80:500-509, DOI:10.1002/jwmg.1030.

Diller, L, K Hamm, D Early, D Lamphear, K Dugger, C Yackulic, C Schwarz, P. Carlson, and T McDonald. 2016. Demographic response of northern spotted owls to barred owl removal in coastal northern California. Journal of Wildlife Management 80:691-707, DOI: 10.1002/jwmg.1046.

Dugger, KM, ED Forsman, AB Franklin, RJ Davis, GC White, CJ Schwarz, KP Burnham, JD Nichols, JE Hines, CB Yackulic, PF Doherty, Jr., L Bailey, DA Clark, SH Ackers, LS Andrews, B Augustine, BL Biswell, J Blakesley, PC Carlson, MJ Clement, LV Diller, EM Glenn, A Green, SA Gremel, DR Herter, J. M Higley, J Hobson, RB Horn, KP Huyvaert, C McCafferty, T McDonald, K McDonnell, GS Olson, JA Reid, J Rockweit, V Ruiz, J Saenz, and SG Sovern. 2016. The effects of habitat, climate and barred owls on long-term demography of northern spotted owls. The Condor: Ornithological Applications 118:57–116, DOI: 10.1650/CONDOR-15-24.1


Wiens, JD, KM Dugger, KE Lewicki, and DC Simon. 2015. Effects of experimental removal of barred owls on population demography of northern spotted owls in Washington and Oregon – 2015 Progress Report. U.S. Geological Survey Open-File Report 2016-1041, 16 p. http://dx.doi.org/10.3133/ofr20161041

Kroll, AJ, TS Garcia, J Jones, KM Dugger, B Murden, J Johnson, S Peterman, B Brintz, and M Rochelle. 2015. Evaluating multi-level models to assess occupancy state responses of plethodontid salamanders. PLoS ONE 10(11): e0142903. doi:10.1371/journal.pone.0142903.

Whitehead, AL, PO’B Lyver, G Ballard, K Barton, BJ Karl, KM Dugger, S Jennings, A Lescroel, PR Wilson, and DG Ainley. 2015. Factors driving Adelie penguin chick size, mass and condition at colonies of different sizes in the southern Ross Sea. Marine Ecology Progress Series 523:199-213.

Sovern, SG, ED Forsman, KM Dugger, and M Taylor. 2015. Habitat use by northern spotted owls during natal dispersal. Journal of Wildlife Management 79:254-262.

Schreck, CB, DD Roby, KM Dugger, and J Peterson. 2015. Meeting cooperator needs: examples from the Oregon Cooperative Fish and Wildlife Research Unit. Proceeding of the North American Wildlife and Natural Resources Conference. Omaha, NE.


Dugger, KM, G Ballard, DG Ainley, PO’B Lyver, and C Schine. 2014. Adélie penguins coping with environmental change: results from a natural experiment at the edge of their breeding range. Frontiers in Ecology and Evolution 2:68. Doi:10.3389/fevo.2014.00068.

Ackers, SH, RJ Davis, KA Olsen, and KM Dugger. 2014. The evolution of mapping habitat for northern spotted owls (Strix occidentalis caurina): a comparison of photo-interpreted, Landsat, and LiDAR based habitat maps. Remote Sensing of Environment 156:361-373.

Kennedy, PL, AM Bartuszevige, M Houle, AB Humphrey, KM Dugger and J Williams. 2014. Long-term dynamics in buteo hawk territory occupancy on a privately-managed bunchgrass prairie in Northeast Oregon. The Condor: Ornithological Applications 116:435-445.


LaRue, MA, DG Ainley, M Swanson, KM Dugger, PO’B Lyver, K Barton, and G Ballard.  2013.  Climate change winners: receding ice fields facilitate colony expansion and altered dynamics in an Adélie penguin metapopulation.  PLoS ONE 8(4):e60568.  Doi:10.1371/journal.pone.0060568.

Schilling, JW, KM Dugger, and RG Anthony. 2013. Survival and home-range size of northern spotted owls in southwestern Oregon. Journal of Raptor Research 47:1-14.

Faaborg, J, WJ Arendt, JD Toms, KM Dugger, WA Cox, and M Canals Mora.  2013.  Long-term decline of a winter-resident bird community in Puerto Rico.  Biodiversity and Conservation 22:63-75.  DOI 10.1007/s10531-012-0399-7.


Davis, RJ, KM Dugger, S Mohoric, L Evers, and WC Aney.  2011.  Northwest forest plan–the first 15 years (1994–2008): status and trends of Northern Spotted Owl populations and habitat. USDA Forest Service General Technical Report PNW-GTR-850. 

Dugger, KM, RG Anthony, and LS Andrews.  2011Transient dynamics of invasive competition: barred owls, spotted owls and the demons of competition present.  Ecological Applications 21:2459-2468.

Forsman, ED, RG Anthony, KM Dugger, EM Glenn, AB Franklin, GC White, CJ Schwarz, KP Burnham, DR Anderson, JD Nichols, JE Hines, JB Lint, RJ Davis, SH Ackers, LS Andrews, BL Biswell, PC Carlson, LV Diller, SA Gremel, DR Herter, JM Higley, RB Horn, JA Reid, J Rockweit, J Schaberel, TJ Snetsinger, and SG Sovern.  2011.  Population demography of northern spotted owls.  Studies in Avian Biology 40.

Raphael, MG, GA Falxa, KM Dugger, BM Galleher, D Lynch, SL Miller, SK Nelson, and RD Young. 2011. Northwest Forest Plan—the first 15 years (1994–2008): status and trend of nesting habitat for the marbled murrelet. USDA Forest Service General Technical Report PNW-GTR-848.


Dugger, KM, DG Ainley, P Lyver, K B, and G Ballard. 2010Survival differences and the effect of environmental instability on breeding dispersal in an Adélie penguin meta-population.  Proceedings of the National Academy of Sciences 107:12375-12380.

Ballard, G, DG Ainley, KM Dugger, and N Nur.  2010.  Foraging strategies of Adélie penguins: adjusting body condition to cope with environmental variability.  Marine Ecology Progress Series 405:287-302.

Faaborg, J, TW Sherry, A Anders, KL Bildstein, KM Dugger, S Gauthreaux, P Heglund, K Hobson, DH Johnson, SC Latta, DJ Levey, P Marra, CL Merkord, E Nol, SI Rothstein, TS Sillett, FR Thompson III, and N Warnock.   2010.  Managing migratory landbirds in the New World: do we know enough?  Ecological Applications 20(2):398-410.

Faaborg, J, TW Sherry, A Anders, K Bildstein, KM Dugger, S. Gauthreaux, P. Heglund, K.A. Hobson, DH Johnson, SC Latta, D Levey, P Marra, CL Merkord , E Nol, SI Rothstein, S Sillett, FR Thompson III, and N Warnock.  2010.  Recent advances in understanding migration systems of New World landbirds.  Ecological Monographs 80(1):2-48.

Ainley, DM, KM Dugger, RG Ford, SD Pierce, DC Reese, RD Brodeur, CT Tynan, JA Barth.  2009.  The spatial association of predators and prey at frontal features in the northern California Current: Competition, facilitation, or merely co-occurrence?  Marine Ecology Progress Series 389:271-294.


Dugger, BD, KM Dugger and LH Fredrickson.  2009. Hooded Merganser (Lophodytes cucullatus), The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology; Retrieved from the Birds of North America Online: http://bna.birds.cornell.edu/bna/species/098doi:10.2173/bna.98

Lesroel, A, KM Dugger, G Ballard, and DG Ainley.  2009.  Effects of individual quality, reproductive success and environmental variability on survival of a long-lived seabird.  Journal of Animal Ecology 78:798-806.


Ainley, DG, KM Dugger, V Toniolo, and I Gaffney.  2007.  Cetacean occurrence patterns in the Amundsen and southern Bellingshausen Seas sector, Southern Ocean.  Marine Mammal Science 23:287-305.

Faaborg, J., KM Dugger, and W. J. Arendt.  2007.  Long-term variation of the winter-resident bird community of Guánica Forest, Puerto Rico: lessons for measuring and monitoring species richness.  Journal of Field Ornithology 78:270-278.

Hagar, JC, KM Dugger, and EE Starkey.  2007.  Diet of Wilson’s warblers and distribution of arthropod prey in the understory of Douglas-fir forests.  Wilson Journal of Ornithology 119:533-546.


Ainley, DG, G. Ballard, and KM Dugger.  2006.  Competition among penguins and cetaceans reveals trophic cascades in the Western Ross Sea, AntarcticaEcology 87:2080-2093.

Anthony, RG, ED Forsman, AB Franklin, DR Anderson, KP Burnham, GC White, CJ Schwarz, JD Nichols, JE Hines, GS Olson, SH Ackers, S Andrews, BL Biswell, PC Carlson, LV Diller, KM Dugger, KE Fehring, TL Fleming, RP Gerhardt, SA Gremel, RJ Gutiérrez, PJ Happe, DR Herter, JM Higley, RB Horn, LL Irwin, PJ Loschl, JA Reid, SS Sovern.  2006.  Status and trends in demography of Northern Spotted Owls, 1985-2003.  Wildlife Monographs 163:1-47.

Dugger, KM, G Ballard, DG Ainley, and K Barton.  2006Effects of flipper-bands on apparent survival and foraging behavior of Adélie penguins.  The Auk 123:858-869.

Sheffield, LM, AE Gall, DD Roby, DB Irons, and KM Dugger2006.  Monitoring planktivorious seabird populations: validating surface counts of crevice-nesting Auklets using mark-resighting.  Canadian Journal of Zoology 84:846-854.


Ainley, DG, G Ballard, BJ Karl, and KM Dugger2005.  Leopard seal predation rates at penguin colonies of different size.  Antarctic Science 17:335-340.

Dugger, KM, FF Wagner, RG Anthony, GS Olson.  2005The relationship between habitat characteristics and demographic performance of Northern Spotted Owls in southern Oregon.  Condor 107:865-880.

Olson, GS, RG Anthony, ED Forsman, SH Ackers, PJ Loschl, JA Reid, KM Dugger, EM Glenn, and WJ Ripple.  2005.  Modeling of site occupancy dynamics of northern spotted owls in Oregon, including the effects of barred owl encroachment.  Journal of Wildlife Management 69:918-932.


Dugger, KM, J Faaborg, WJ Arendt, and KA Hobson.  2004Understanding survival and abundance of overwintering warblers: does rainfall matter?  Condor 106:744-760.

Faaborg, J, WJ Arendt, and KM Dugger2004.  Bird population studies in Puerto Rico using mist nets: general patterns and comparisons with point counts.  Studies in Avian Biology 29:144-150.

McCreedy, CD, KA Reynolds, CM Basile, MC Nicholson, KM Dugger, ME Gross, GM Mohr, Jr.  2004.  Terrestrial animal species in the Hoosier-Shawnee ecological assessment area.  General Technical Report NC-244.  St. Paul, MN: U.S. Department. of Agriculture, Forest Service, North Central Research Station.  267 pp.

2003 and earlier

Dugger, BD, and KM Dugger2003.  Long-billed Curlew.  Pages 224-226 in Birds of Oregon: A general reference.  D. B. Marshall, M. G. Hunter, and A. L. Contreras, eds. Oregon State University Press, Corvallis, OR.

Dugger, BD and KM Dugger2002.  The long-billed curlew.  In The Birds of North America, No. 628 (A. Poole, P. Stettenheim, and F. Gill, eds.).  Philadelphia:  The Academy of Natural Sciences; Washington, DC; The American Ornithologists’ Union.

Dugger, KM, MR Ryan, DL Galat, RR Renken, JW Smith.  2002Reproductive success of the interior least tern (Sterna antillarum) in relation to hydrology on the Lower Mississippi River.  River Research and Application.  18:97-105.

Dugger, KM, J Faaborg, and WJ Arendt.  2000.  Rainfall correlates of bird populations and survival rates in a Puerto Rican dry forest.  Bird Populations.  5:11-27.

Dugger, KM, MR Ryan, and RR Renken.  2000Survival rates of least tern chicks on the Lower Mississippi River.  Journal of Field Ornithology 71:330-338.

Faaborg, J, Arendt, WJ, and KM Dugger2000.  The Guánica, Puerto Rico, bird monitoring project.  Bird Populations 5:102-111.

Dugger, KM, BD Dugger, and LH Fredrickson.  1999Annual survival rates of female hooded mergansers and wood ducks in the southeast.  Wilson Bulletin 111:1-6.

Faaborg, J, KM Dugger, W Arendt, M Baltz and B Woodworth.  1997.  Evidence for precipitous declines in the Puerto Rican Vireo of Guánica Forest. Wilson Bulletin 109:195-202.

Dugger, BD, KM Dugger, and LH Fredrickson.  1994.  The hooded merganser.  In The Birds of North America, No. 98 (A. Poole, P. Stettenheim, and F. Gill, eds.). Philadelphia: The Academy of Natural Sciences; Washington, DC; The American Ornithologists’ Union.

Dugger, KM, and LH Fredrickson.  1992.  Life history and habitat needs of the wood duck.  Waterfowl Management Handbook, Fish and Wildlife Leaflet13.1.6.