• Ramirez, MD, L Avens, JA Seminoff, LR Goshe, and SS Heppell. 2017. Growth dynamics of loggerhead sea turtles undergoing ontogenetic habitat shifts. Oecologia 183: 1087-1099. DOI: http://dx.doi.org/10.1007/s00442-017-3832-5
  • Egerton JP, Johnson AF, Le Vay L, McCoy CM, Semmens BX, Heppell SA, Turner JR. 2017. Hydroacoustics for the discovery and quantification of Nassau grouper (Epinephelus striatus) spawning aggregations. Coral Reefs 2017:1-2. DOI: http://dx.doi.org/10.1007/s00338-017-1542-4
  • Schwartzkopf, BS, and JH Cowan Jr. 2017. Seasonal and sex differences in energy reserves of red snapper Lutjanus campechanus on natural and artificial reefs in the northwestern Gulf of Mexico. Fisheries Science 83: 13-22. DOI:10.1007/s12562-016-1037-1


  • Gračan, R, SA Heppell, G Lacković, and B Lazar. 2016. Age and growth dynamics of spiny dogfish, Squalus acanthias, in the Adriatic Sea (Eastern Mediterranean Sea). Marine and Freshwater Research 67: 357–367. DOI: http://dx.doi.org/10.1071/MF14116
  • Casale, P, and SS Heppell. 2016. How much sea turtle by-catch is too much? A stationary age distribution model for simulating population abundance and potential biological removal in the Mediterranean. Endangered Species Research 29:239-254. DOI: http://dx.doi.org/10.3354/esr00714
  • Montero-Styles, JT, RO Martinez-Rincon, SS Heppell, M Hall and M Ewal. 2016. Characterizing Environmental and Spatial variables associated with incidental catch of Olive Ridley (Lepidochelys olivacea) in the Eastern Tropical Pacific purse-seine fishery. Fisheries Oceanography 25(1):1-14. DOI: http://dx.doi.org/10.1111/fog.12130


  • Archer, SK, JE Allgeier, BX Semmens, SA Heppell, CV Pattengill-Semmens, AD Rosemond, PG Bush, CM McCoy, BC Johnson, and CA Layman. 2015. Hot moments in spawning aggregations: Implications for ecosystem-scale nutrient cycling. Coral Reefs 34:19–23. DOI: http://dx.doi.org/10.1007/s00338-014-1208-4
  • Páez, VP, A Lipman, BC Bock, and SS Heppell. 2015. A plea to redirect and evaluate conservation programs for South America’s Podocnemidid river turtles. Chelonian Conservation and Biology, 14(2), 205-216. DOI: http://dx.doi.org/10.2744/CCB-1122.1
  • Turner Tomaszewicz, C, JA Seminoff, MD Ramirez, and CM Kurle. 2015. Effects of demineralization on the stable isotope analysis of bone samples. Rapid Communications in Mass Spectrometry, 29: 1879-1888. DOI: http://dx.doi.org/10.1002/rcm.7295
  • Ramirez, MD, L Avens, JA Seminoff, LR Goshe, and SS Heppell. 2015. Patterns of loggerhead turtle ontogenetic shifts revealed through isotopic analysis of annual skeletal growth increments. Ecosphere 6(11): art244. DOI: http://dx.doi.org/10.1890/ES15-00255.1
  • Páez, VP, BC Bock, PA Espinal-Garcıa, BH Rendon-Valencia, D Alzate-Estrada, VM Cartagena-Otalvaro, and SS Heppell. 2015. Life History and Demographic Characteristics of the Magdalena River Turtle (Podocnemis lewyana): Implications for Management. Copeia 103(4):1058-1074. DOI: http://dx.doi.org/10.1643/CE-14-191
  • Piacenza SE, AK Barner, CE Benkwitt, KS Boersma, EB Cerny-Chipman, KE Ingeman, TL Kindinger, JD Lee, AJ Lindsley, JN Reimer, JC Rowe, C Shen, KA Thompson, LL Thurman, and SS Heppell. 2015. Patterns and variation in benthic biodiversity in a large marine ecosystem. PLoS ONE 10(8): e0135135. DOI: http://dx.doi.org/10.1371/journal.pone.0135135
  • Piacenza SE, LL Thurman, AK Barner, CE Benkwitt, KS Boersma, EB Cerny-Chipman, KE Ingeman, TL Kindinger, AJ Lindsley, J Nelson, JN Reimer, JC Rowe, C Shen, KA Thompson, and SS Heppell. 2015. Evaluating temporal consistency in marine biodiversity hotspots. PLoS ONE 10(7): e0133301. DOI: http://dx.doi.org/10.1371/journal.pone.0133301
  • Easton RR, SS Heppell, and RW Hannah. 2015. Quantification of habitat and community relationships among nearshore temperate fishes through analysis of drop camera video. Marine and Coastal Fisheries 7:87-102. DOI: http://dx.doi.org/10.1080/19425120.2015.1007184


  • Jones, NM, BA Hoover, SA Heppell, and KJ Kuletz. 2014. A cross-shelf gradient in δ15N stable isotope values of krill and pollock Indicates seabird foraging patterns in the Bering Sea. Deep Sea Research Part II Topical Studies in Oceanography 109:241-250. DOI: http://dx.doi.org/10.1016/j.dsr2.2014.04.008
  • Jackson, AM, BX Semmens, Y Sadovy de Mitcheson, RS Nemeth, SA Heppell, et al. (2014) Population structure and phylogeography in Nassau Grouper (Epinephelus striatus), a mass-aggregating marine fish. PLoS ONE 9(5): e97508. DOI: http://dx.doi.org/10.1371/journal.pone.0097508
  • Grüss, A, J Robinson, SS Heppell, SA Heppell, and BX Semmens. 2014. Conservation and fisheries effects of spawning aggregation marine protected areas: What we know, where we should go, and what we need to get there. ICES Journal of Marine Science 71(7):1515-1534. DOI: https://doi.org/10.1093/icesjms/fsu038
  • Paredes, R, RA Orben, RM Suryan, DB Irons, DD Roby, AMA Harding, RC Young, KJ Benoit-Bird, C Ladd, H Renner, SA Heppell, RA Phillips, and A Kitaysky. 2014. Foraging responses of Black- Legged Kittiwakes to prolonged food shortages around colonies on the Bering Sea shelf. PLoS ONE 9(3): e92520. DOI: http://dx.doi.org/10.1371/journal.pone.0092520
  • Melnychuk, M, TE Essington, TA Branch, SS Heppell, OP Jensen, JS Link, SJD Martell, AM Parma, and ADM Smith. 2014. Which design elements of individual quota fisheries help to achieve management objectives? Fish and Fisheries 17:126-142. DOI: http://dx.doi.org/10.1111/faf.12094
  • Cianelli, L, M Hunsicker, A Beaudreau, K Bailey, L Crowder, C Finley, C Webb, J Reynolds, K Sagmiller, JM Anderies, D Hawthorne, J Parrish, S Heppell, F Conway, and P Chigbu. 2014. Transdisciplinary graduate education in marine resource science and management. ICES Journal of Marine Science 71(5):1047-1051. DOI: http://dx.doi.org/10.1093/icesjms/fsu067
  • Arnold, LM, and SS Heppell. 2014. Testing the robustness of data-poor assessment methods to uncertainty in catch and biology: a retrospective approach. ICES Journal of Marine Science. 72:243-250. DOI: http://dx.doi.org/10.1093/icesjms/fsu077
  • Pikitch EK, KJ Rountos, TE Essington, C Santora, D Pauly, R Watson, UR Sumaila, PD Boersma, IL Boyd, DO Conover, P Cury, SS Heppell, ED Houde, M Mangel, E Plaganyi-Lloyd, K Sainsbury, RS Steneck, TM Geers, N Gownaris, SB Munch. 2014. The global contribution of forage fish to marine fisheries and ecosystems. Fish and Fisheries 15(1):43-64. DOI: http://dx.doi.org/10.1111/faf.12004
  • Senko, J, ER White, SS Heppell, and LR Gerber. 2014. Comparing bycatch mitigation strategies for vulnerable marine megafauna. Animal Conservation. 17(1):5-18. DOI: http://dx.doi.org/10.1111/acv.1205
  • Thompson, KA, SS Heppell, and GG Thompson. 2014. The effects of temperature and predator densities on the consumption of walleye pollock (Gadus chalcogrammus) by three groundfish in the Gulf of Alaska. Canadian Journal of Fisheries and Aquatic Sciences 71(8):1123-1133. DOI: http://dx.doi.org/10.1139/cjfas-2013-0260


  • Kammerer, BD, and SA Heppell. 2013. The effects of semi-chronic thermal stress on physiological indicators in steelhead Oncorhynchus mykiss. Transactions of the American Fisheries Society 142: 1299-1307. DOI: http://dx.doi.org/10.1080/00028487.2013.806349
  • Benoit-Bird, KJ, NM McIntosh, and SA Heppell. 2013. Nested scales of spatial heterogeneity of juvenile walleye pollock (Theragra chalcogramma) in the southeastern Bering Sea. Marine Ecology Progress Series 484: 219-238. DOI: http://dx.doi.org/10.3354/meps10319
  • Benoit-Bird, KJ, BC Battaile, SA Heppell, B Hoover, D Irons, N Jones, KJ Kuletz, CA Nordstrom, R Paredes, RM Suryan, CM Waluk, and AW Trites. 2013. Prey patch patterns predict habitat use by top marine predators with diverse foraging strategies. PLoS ONE 8: e53348. DOI: http://dx.doi.org/110.1371/journal.pone.0053348
  • Barcelo, C, A Domingo, P Miller, L Ortega, B Giffoni, G Sales, L McNaughton, M Marcovaldi, SS Heppell, and Y Swimmer. 2013. High-use areas, seasonal movements and dive patterns of juvenile loggerhead sea turtles in the Southwestern Atlantic Ocean. Marine Ecology Progress Series 479:235-250. DOI: http://dx.doi.org/10.3354/meps10222
  • Chesney,TA, SS Heppell, J Montero, and J Graham. 2013. Interannual variability of Humboldt squid (Dosidicus gigas) off Oregon and Southern Washington. Cal-COFI Reports 54:180-191.
  • Cook, MT, SS Heppell, and TS Garcia. 2013. Invasive bullfrog larvae lack developmental plasticity to changing hydroperiod. Journal of Wildlife Management 77: 655–662. DOI: http://dx.doi.org/10.1002/jwmg.509
  • Moore, J, K Curtis, R Lewison, P Dillingham, J Cope, S Fordham, SS Heppell, S Pardo, C Simpfendorfer, and G Tuck. 2013. Evaluating sustainability of fisheries bycatch mortality for marine megafauna: a review of conservation reference points for data-limited populations. Environmental Conservation 40(4):329-344. DOI: http://dx.doi.org/10.1017/S037689291300012X
  • Smith, WD, J Miller, and SS Heppell. 2013. Elemental markers in elasmobranchs: the effects of environmental history and growth on vertebral chemistry. PloS ONE 8(10): e62423. doi:10.1371/journal.pone.0062423. DOI: http://dx.doi.org/10.1371/journal.pone.0062423